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First, that the GSH binding site(s) on the structure of the capsid is completely different than the locations of the BSO resistant mutations

doi: 10.1074/jbc.M116.731216 82 DonskovFHoklandMMarcussenNTorp MadsenHHvon der MaaseH

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In all of these conditions, the glucose levels were decreased when compared with the wild type control animals, with the lowest glucose levels being associated with the simultaneous deletion of the three isoforms

Mechanistically, these data, together with observed decrease in proton leakage, indicated profound changes in mitochondrial function leading to energy depletion